Scotland lose Tierney for Euro 2020 opener
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Tierney has been one of the most consistent performers during Scotland's improvement under Steve Clarke over the past two years, deputising at centre-back to allow captain Andy Robertson to play in his natural role as a left-back.
Elsewhere, Clarke has kept faith with a number of key players from playoff wins over Israel and Serbia to finally make it back to a major tournament.
Stuart Armstrong and Ryan Christie are preferred to Chelsea's Billy Gilmour and Southampton striker Che Adams.
The Czechs are without suspended defender Ondrej Kudela after his 10-match ban for racially abusing Rangers midfielder Glen Kamara during a Europa League last 16 clash with Slavia Prague.
West Ham duo Vladimir Coufal and Tomas Soucek start, while Bayer Leverkusen's Patrik Schick leads the line up front.
Scotland (5-3-2)
David Marshall; Stephen O'Donnell, Jack Hendry, Liam Cooper, Grant Hanley, Andy Robertson (capt); Scott McTominay, Stuart Armstrong, John McGinn; Lyndon Dykes, Ryan Christie
Coach: Steve Clarke
Czech Republic (4-2-3-1)
Tomas Vaclik; Vladimir Coufal, Ondrej Celustka, Tomas Kalas, Jan Boril; Tomas Soucek, Alex Kral; Lukas Masopust, Vladimir Darida (capt), Jakub Jankto; Patrik Schick
Coach: Jaroslav Silhavy
Referee: Daniel Siebert (GER)